Spesies Baru dan Spesies Rekam Baru Serangga Kutu Putih (Hemiptera: Pseudococcidae) di Indonesia

Authors

Agustin Zarkani, dkk.
Universitas Bengkulu

Synopsis

Kutu putih atau mealybug (Hemiptera: Coccomorpha: Pseudococcidae) adalah salah satu
kelompok serangga pencucuk penghisap (sucking insects) dari kelompok kutu sisik (scale insects)
yang keberadaanya sangat penting di ekosistem pertanian dengan peran dominannya sebagai hama
tanaman di seluruh dunia. Di Indonesia, dalam rentang tahun 2020 hingga 2024 telah ditemukan
4 spesies baru kutu putih dunia serta 9 spesies rekam baru untuk Indonesia. Empat spesies baru
dunia tersebut yaitu Chrorizococcus ozeri Zarkani & Kaydan, Dysmicoccus zeynepae Zarkani &
Kaydan, Planococcus bengkuluensis Zarkani & Kaydan, dan Pseudococcus iffahae Zarkani &
Kaydan. Adapun 9 spesies rekam baru Indonesia yaitu Ferrisia dasylirii (Cockerell), Planococcus
bendovi Williams, Rastrococcus tropicasiaticus Williams, Coccidohyistrix insolita (Green),
Dysmicoccus arachidis Williams, Dysmicoccus carens Williams, Paracoccus tripurae Williams,
Planococcus bagmaticus Williams dan Pseudococcus leptotrichotus Williams. Data lengkap
morfologi semua spesies baru dan rekam baru tersebut dimuat dalam buku ini

References

Arifin, H.S. & Nakagoshi, N. (2011) Landscape ecology and

urban biodiversity in tropical Indonesian cities.

Landscape and Ecological Engineering, 7 (1), 33–43.

https://doi.org/10.1007/s11355-010-0145-9

Ben-Dov, Y. (1994) A systematic catalogue of the mealybugs of

the world (Insecta: Homoptera: Coccoidea:

Pseudococcidae and Putoidae) with data on

geographical distribution, host plants, biology and

economic importance. Intercept Limited, Andover,

U.K., 686 pp.

Betrem, J.G. (1937) De morporphologie en systematiek van

enkele vande vooraamste witte-luizensoorten van Java.

Archief voor de Koffiecultuur in Nederlandsch-Indie,

, 1‒118.

Betrem, J.G. (1940) A new Rhizoecus species. Treubia, 17,

‒270.

CABI. (1959) Saccharicoccus sacchari (Ckll.). Distribution

Maps of Insect Pests. Series A Map no. 102, 2 pp.

CABI. (1983) Nipaecoccus viridis (Newstead). Distribution

Maps of Pests, Series A, Agricultural Map no. 446, 2

pp.

CABI. (1995) Planococcus lilacinus (Cockerell). Distribution

Maps of Pests Map no. 101 (1st rev.), 4 pp.

CBD Secretariat (2021) Indonesia–Country Profile:

Biodiversity Facts.

https://www.cbd.int/countries/profile/default.shtml?coun

try=id#facts (accessed 22 March 2021). Gavrilov-Zimin

IA (2013) New scale insects (Homoptera: Coccinea)

from Sulawesi and New Guinea, with some other

additions to the Indonesian fauna. Tropical Zoology 26;

‒86. https://doi.org/10.1080/03946975.2013.807570

Charles, J.G. (1981) Distribution and life history of the long

tailed mealy bug, Pseudococcus longispinus

(Homoptera: Pseudococcidae), in Auckland vineyards.

New Zealand Journal of Zoology, 8: 285-293.

Cox, J.M. (1989) The mealybug

genus Planococcus (Homoptera: Pseudococcidae).

Bulletin British Museum (Natural History). Entomology,

(1), 1‒78.

Daane, K.M., Almeida, R.P.P., Bell, V.A., Walker, J.T.S.,

Botton, M., Fallahzadeh, M., Mani, M., Miano, J.L.,

Sforza, R., Walton, V.M. & Zaviezo, T. 2012. Biology

and management of mealybugs in vineyards. In

Bostanian, N.J., Vincent, C. & Isaacs, R. (eds.).

Arthropod Management in Vineyards, pp. 271–307.

Dordrecht: Springer Netherlands Press.

Dammerman, K.W. (1929) The agricultural zoology of the

Malay Archipelago. The animals injurious and

beneficial to agriculture, horticulture and forestry in the

Malay Peninsula, the Dutch East Indies and

Philippines. J.H. de Bussy I‒XI, Amsterdam,

Netherlands, 473 pp.

Danzig, E.M. & Gavrilov, I.A. 2010. Mealybugs of the genera

Planococcus Ferris and Crisicoccus Ferris

(Sternorrhyncha: Pseudococcidae) of Russia and

adjacent countries. Zoosystematica Rossica 19(1): 39-

Dewi, N., Wijayanto, N. & Gusmaini. 2017. Dimension growth

of Azadirachta excelsa and Phyllanthus spp. in

agroforestry system. Biodiversitas 18(2): 494-499.

Francardi, V. & Covassi, M. 1992: Note bio-ecologishe sul

Planococcus novae (Nasonov) dannoso a Juniperus spp.

in Toscana (Homoptera: Pseudococcidae). Redia 75: 1–

Franco, J.C., Zada, A. & Mendel, Z. 2009. Novel approaches

for the management of mealybug pests. In Ishaaya, I. &

Horowitz, A.R. (eds.). Biorational Control of Arthropod

Pests, pp. 233–278. New York: Springer

Science + Business Media.

Ganguli, R.N. & Ghosh, M.R. 1964. Coccoids of Tripura.

Indian Journal of Entomology 26: 358-359.

Garcia-Morales, M., Denno, B.D, Miller, D.R., Miller, G.L.,

Ben-Dov, Y. & Hardy, N.B. 2016. ScaleNet: A

literature-based model of scale insect biology and

systematics. http://scaleFErrisnet.info [12 Juli 2022].

Gavrilov-Zimin, I.A. (2012) A contribution to the taxonomy,

cytogenetics and reproductive biology of the genus

Aclerda Signoret (Homoptera, Coccinea). Comparative

Cytogenetics, 6(4), 389‒395.

https://doi.org/10.3897/compcytogen.v6i4.4320

Gavrilov-Zimin, I.A. (2013) New scale insects (Homoptera:

Coccinea) from Sulawesi and New Guinea, with some

other additions to the Indonesian fauna. Tropical

Zoology, 26 (2), 64‒86.

https://doi.org/10.1080/03946975.2013.807570

Gavrilov-Zimin, I.A. (2016) Cytogenetic and taxonomic studies

of some legless mealybugs (Homoptera, Coccinea,

Pseudococcidae). Comparative Cytogenetics, 10 (4),

‒601.

https://doi.org/10.3897/compcytogen.v10i4.10503

Gavrilov-Zimin, I.A. (2017) Taxonomic and faunistic notes on

scale insects (Homoptera: Coccinea) of Bali, Flores and

New Guinea (Indonesia). Biodiversity, Biogeography

and Nature Conservation in Wallacea and New Guinea.

Entomological Society of Latvia Salaspils, 3, 141‒149.

Gavrilov-Zimin, I.A. (2018a) Ontogenesis, morphology and

higher classification of archacococcids (Homoptera:

Coccinea: Orthezoiidea). Zoosystematica Rossica, 2, 1‒

https://doi.org/10.31610/zsr/2018.supl.2.1

Gavrilov-Zimin, I.A. (2018b) First illustration of chromosomes

and genetic system of Lecanodiaspidinae (Homoptera:

Coccinea: Asterolecaniidae s.l.). Comparative

Cytogenetics, 12 (3), 439‒443.

https://doi.org/10.3897/CompCytogen.v12i3.29648

Gavrilov-Zimin, I.A. (2019) New Asterolecaniidae s.l.

(Homoptera: Coccinea) from different regions of the

world and some distributional records. Tropical

Zoology, 32, 142‒154.

https://doi.org/10.1080/03946975.2019.1641673

Gavrilov-Zimin, I.A. (2020) Chromosomal and reproductive

features of some Oriental and Australasian scale insects

(Homoptera, Coccinea). Comparative Cytogenetics, 14

(3), 339–352.

https://doi.org/10.3897/CompCytogen.v14i3.53367

Gavrilov-Zimin, I.A. (2021) New and poorly known giant scale

insects (Homoptera: Coccinea: Margarodidae s. lat.)

from the Oriental region with taxonomic and

nomenclatural notes on the subfamily

Monophlebinae. European Journal of Taxonomy 746

(1), 50–61.

https://doi.org/10.5852/ejt.2021.746.1317

Giliomee, J.H. (1966) A list of South African scale insects

(Homoptera: Coccoidea) in the collection of the British

Museum (Natural History) with information on their

host plants and distribution. Annals of the University of

Stellenbosch (Section A), 41, 411‒428.

Ginting, S., Depari, E.K., Nadrawati & Zarkani, A. 2020. The

attack of Rastrococcus sp. (Hemiptera: Pseudococcidae)

on Dysoxylum mollissimum Blume in campus forest of

Bengkulu University. Jurnal Hama dan Penyakit

Tumbuhan Tropika 20(1): 54-60.

Green, E.E. (1922) The Coccidae of Ceylon, Part V. Dulau &

Co., London, U.K., 345‒472.

Green, E.E. (1930) Fauna Sumatrensis (Bijdrag Nr. 65).

Coccidae. Tijdschrift voor Entomologie, 73, 279‒297.

Gullan, P.J., Downie, D.A. & Steffan, S.A. (2003) A new pest

species of the mealybug genus Ferrisia Fullaway

(Hemiptera: Pseudococcidae) from the United States.

Annals of the Entomological Society of America 96 (6):

–737.

Gullan, P.J. & Martin, J.H. 2009. Sternorrhyncha (Jumping

plant-lice, whiteflies, aphids, and scale insects). In Resh,

V.H. & Cardé, R.T. (eds.) Encyclopedia of Insects. 2

nd

Edition, pp. 957–967. San Diego: Elsevier.

Gullan, P.J., M.B. Kaydan and N.B. Hardy. ( 2010)

Molecular phylogeny and species recognition in the

mealybug genus Ferrisia Fullaway (Hemiptera:

Pseudococcidae). Systematic Entomology 35: 329–339

Hodgson, C.J. (1994) The Scale Insect Family Coccidae: An

Identification Manual to Genera. UK: CAB

International Wallingford.

Kalshoven, L.G.E. (1981) Coccoidea. Pests of crops in

Indonesia. Ichtiar Baru, Jakarta, Indonesia, 701 pp.

Kaydan M.B. and P.J. Gullan. ( 2012) A taxonomic

revision of the mealybug genus Ferrisia Fullaway

(Hemiptera: Pseudococcidae), with descriptions of

eight new species and a new genus. Monograph.

Zootaxa 3543: 1–65.

Kaydan M.B., F. Kozar and C. Hodgson. (2015) A review of

the phylogeny of Palaearctic mealybugs (Hemiptera:

Coccomorpha: Pseudococcidae). Arthropod Systematic

and Phylogeny 73(1): 175–195

Keuchenius, P.E. (1915) Onderzoekingen en beschouwingen

over eenige schadlijke schildluizen van de

Koffiekultuur op Java. Mededeelingen van het

Besoekisch Proefstation. Djember, 16, 1‒65.

Kottek, M., Grieser, J., Beck, C., Rudolf, B. & Rubel, F. (2006)

World map of the Köppen-Geiger climate classification

updated. Meteorologische Zeitschrift, 15 (3), 259-263.

https://doi.org/10.1127/0941-2948/2006/0130

Kosztarab, M. & Kozár, F. (1988). Scale Insects of Central

Europe. Akadémiai Kiadó, Budapest. 600 pp.

Krantz GW, Walter DE. 2009. A Manual Acarology. Third

Edition. Texas (US): Texas Tech University Press.

Lohman, D., Bruyn, M., Page, T., Rintelen, K., Hall, R., Ng,

P.L., Shih, H., Carvalho, G. & Rintelen, T. (2011)

Biogeography of the Indo-Australian Archipelago.

Annual Review of Ecology, Evolution, and Systematics,

(1), 205‒226.

Magallón-Servín, P., López-Vela, M., Pedraza-Mirafuentes, M.

& Servín-Villegas, R. (2019) Ferrisia dasylirii

(Cockerell, 1896), a Potential pest of commercial

plantations of Salicornia bigelovii (Torr.) at Baja

California Sur, Mexico. Southwestern Entomologist 44

(4): 861–866.

https://doi.org/10.1146/annurev-ecolsys-102710-145001

McNeely, J.A., Miller; K.R., Reid, W.V., Mittermeier, R.A. &

Werner, T.B. (1990) Conserving the World's Biological

Diversity. IUCN, Gland, Switzerland, 193 pp.

Muniappan, R., Shepard, B.M., Watson, G.W., Carner, G.R.,

Sartiami, D., Rauf, A., Xiong, J.J. & Hammig, M.D.

(2008) First report of the Papaya Mealybug, Paracoccus

marginatus (Hemiptera: Pseudococcidae), in Indonesia

and India. Journal of Agricultural and Urban

Entomology 25 (1), 37‒40.

https://doi.org/10.3954/1523-5475-25.1.37

Muniappan, R., Shepard, B.M., Watson, G.W., Carner, G.R.,

Rauf, A., Sartiami, D., Hidayat, P, Afun, J.V.K.,

Goergen, G. & Ziaur Rahman, A.K.M. (2011) New

records of invasive insects (Hemiptera: Sternorrhyncha)

in Southeast Asia and West Africa. Journal of

Agricultural and Urban Entomology, 26 (4), 167‒174.

https://doi.org/10.3954/1523-5475-26.4.167

Muniappan, R., Watson, G.W., Evans, G.A., Rauf, A. & von

Ellenrieder, N. (2012) Cycad Aulacaspis scale, a newly

introduced insect pest in Indonesia. HAYATI Journal of

Biosciences, 19 (3), 110‒114.

https://doi.org/10.4308/hjb.19.3.110

Nabil, A.N. & Hegab, M.A.M. (2019) Impact of some weather

factors on the population density of Phenacoccus

solenopsis Tinsley and its natural enemies. Egypt. Acad.

J. Biolog. Sci., 12(2): 99– 108.

Newstead, R. (1908) On a collection of Coccidae and other

insects affecting some cultivated and wild plants in Java

and in Tropical Western Africa. Journal of Economic

Biology. London, 3, 33‒42.

Nurkomar, I., Trisnawati, D.W. & Aisyah, S.N. (2021) Pest

insects in cassava plantations intercrop with different

plants at Gunungkidul, Yogyakarta, Indonesia.

Serangga 26(1):1-12.

Pacheco da Silva V. C., Kaydan M.B., Da Silva-Torres C. S.

A. and Torres J. B.,2019. Mealybug species

(Hemiptera: Coccomorpha: Pseudococcidae) on soursop

and sugar apple (Annonaceae) in North-East Brazil,

with description of a new species of Pseudococcus

Westwood. Zootaxa 4604 (3): 525–538

Reyne, A. (1954a) Hippeococcus a new genus of

Pseudococcidae from Java with peculiar habits.

Zoologische Mededelingen Uitgegeven door het

Rizksmuseum van Natuurlijke Historie te Leiden, 32,

‒257.

Reyne, A. (1957) Walkeriana tosariensis, a new monophlebine

coccid from Casuarina junghununiana Miq. in eastJava. Beaufortia, Amsterdam, 6, 115‒145.

Reyne, A. (1961) Scale insects from Dutch New Guinea.

Beaufortia, Amsterdam, 8, 121‒167.

Reyne, A. (1965a) Observations on some Indonesian scale

insects. Tijdschrift voor Entomologie, Amsterdam, 108,

‒188.

Riedel, A. & Narakusumo R.P. (2019) One hundred and three

new species of Trigonopterus weevils from Sulawesi.

ZooKeys, 7 (828), 1‒153.

https://doi.org/10.3897/zookeys.828.32200

Rizwan, M., Raza, A. B. M., Majeed, M. Z., & Arshad, M.

Population Dynamics of Mealybug Drosicha

mangiferae (Green) (Hemiptera: Pseudococcidae) in

Citrus Orchards of District

Sartiami, D., Watson, G.W., Mohamad Roff, M.N. & Idris,

A.B. (2016) New Indonesian country records and

species information for mealybugs (Hemiptera

Pseudococcidae) in Wirjati’s historic collection. Redia

XCIX, 155‒161. https://doi.org/10.19263/Redia99.16.20

Sartiami, D., Sosromarsono, S., Buchori, D. & Suryobroto, B.

(1999) Keragaman spesies kutu putih pada tanaman

buah-buahan di daerah Bogor. Prosiding Seminar

Nasional Perhimpunan Entomologi Indonesia (PEI),

, hlm. 429–435.

Sartiami, D., Watson, G.W., Mohamad, Roff, M.N. & Idris,

A.B. (2016) Mealybugs (Hemiptera: Coccomorpha:

Pseudococcidae) attacking Hibiscus rosa-sinensis L. in

Malaysia, with two new country records. AIP Conference

Proceedings 1784(1): 1-7.

Sartiami, D., Watson, G.W., Roff, M.N.M, & Ghani, I.A.

(2017) A taxonomic update of Takahashi’s historic

collection of mealybugs (Hemiptera: Pseudococcidae)

from Malaysia and Singapore. Serangga 22 (2): 91-114.

Sartiami, D., Adisuseno, W.A., Rachmatika, N., Anwa, R. &

Watson, G.W. (2022) Coccidohystrix insolita (green)

(Hemiptera: Pseudococcidae) from Indonesia and its

note on the three solanaceous plants. Serangga 27 (3):

-171.

Shah, T.N., Ahmed, A.M., & Memon, N. (2015) Population

dynamics of cotton mealybug, Phenacoccus solepnosis

Tinsely in three talukas of district Sanghar (Sindh). J.

Entomol. Zool. Studies, 3(5): 162-167.

Shothwood, T.R.E. (1984) Ecological Methods, 2nd edition.

Chapman and Hall, New York

Supartha, I.W., Yudha, I.K.W., Wiradana, P.A. & Susila, I.W.

(2020) Response of parasitoids to invasive pest

Phenacoccus manihoti Matile-Ferrero (Hemiptera:

Pseudococcidae) on cassava crop in Bali, Indonesia.

Biodiversitas, 21 (10), 4543‒4349.

https://doi.org/10.13057/biodiv/d211011

Takahashi, R. (1950) Some species of Coccidae from the

Riouw Is - Part I. (Homoptera). Insecta Matsumurana,

, 65‒72.

Telli, S. & Yiğit, A. (2019) Population fuctuations

of the citriculus mealybug, Pseudococcus cryptus

Hempel (Hemiptera: Pseudococcidae), in citrus

orchards of Samandağ, Hatay, Turkey. Journal of Plant

Diseases and Protection, 126: 421–426.

Upton, M.S. (1991) Methods for Collecting, Preserving and

Studying Insects, and Allied Forms. Australian

Entomological Society, Miscellaneous Publication No. 3.

Brisb ane (AU): Australian Entomological Society.

Van Heurn, W.C. (1923) The insect pests of rice in Java.

Mededeelingen van het Intituut vor Plantenziekten, 61,

‒43.

Varshney, R.K. 1992. A check list of the scale insects and

mealybugs of South Asia. Part-1. Records of Zoological

Survey of India Occasional Paper 139: 1-152.

Walker AK, Crosby TK. (1988) The Preparation and Curation

of Insects. DSIR Information Series 163, SIPC, DSIR.

Wellington (NZ): DSIR.

Watson, G.W., Muniappan, R., Shepard, B.M., Sembel, D.T.,

Xiong, J.J. & Rauf, A. (2014) Sap-sucking insect

records (Hemiptera: Sternorrhyncha and Thysanoptera:

Thripidae) from Indonesia. Florida Entomologist, 97

(4), 1594‒1597.

https://doi.org/10.1653/024.097.0432

Williams, D.J. (1970) The mealybugs (Homoptera, Coccoidea,

Pseudococcidae) of sugar-cane, rice and sorghum.

Bulletin of Entomological Research, 60, 109‒188.

https://doi.org/10.1017/S0007485300034209

Williams, D.J. (1978) The anomalous ant-attended mealybugs

(Homoptera: Pseudococcidae) of South-East Asia.

Bulletin of the British Museum (Natural History)

Entomology, 37, 72 pp.

Williams, D.J. (1985) Australian mealybugs. British Museum

(Natural History), London, U.K., 431 pp.

Williams, D.J. (1989) The mealybug genus Rastrococcus Ferris

(Hemiptera: Pseudococcidae). Systematic Entomology,

(4), 433‒486.

https://doi.org/10.1111/j.1365-3113.1989.tb00298.x

Williams, D.J. (1996) A brief account of the hibiscus mealybug

Maconellicoccus hirsutus (Hemiptera: Pseudococcidae),

a pest of agriculture and horticulture, with descriptions

of two related species from southern Asia. Bulletin of

Entomological Research, 86, 617‒628.

https://doi.org/10.1017/S0007485300039420

Williams, D.J. (2002) IV The mealybug tribe

Allomyrmococcini and its association with herdsman

ants of the genus Dolichoderus in southern Asia. In:

Dill, M., Williams, D.J & Maschwitz, U. Herdsmen ants

and their mealybug partners. Abhandlungen der

Senckenbergischen Naturforschenden Gesellschaft, pp.

‒178.

Williams, D.J. (2004) Mealybugs of southern Asia. The Natural

History Museum, London, U.K. Southdene SDN BHD,

Kuala Lumpur, Malaysia, 896 pp.

Williams, D.J. & Watson, G.W. (1988a) The Scale Insects of

the Tropical South Pacific Region. Pt. 1. The Armoured

Scales (Diaspididae). CAB International, Wallingford,

U.K., 290 pp.

Williams, D.J. & Watson, G.W. (1988b) The Scale Insects of

the Tropical South Pacific Region. Pt. 2. The Mealybugs

(Pseudococcidae). CAB International, Wallingford,

U.K., 260 pp.

Wirjati. (1958) On a small mealybug from a virgin forest. Idea,

Journal of the Entomological Society of Indonesia, 11,

‒17.

Wirjati. (1959) A new species of Trionymus found on rice-roots

in Java. Idea, Journal of the Entomological Society of

Indonesia, 12, 14‒19.

Zarkani, A., Apriyanto, D., Turanli, F. & Kaydan, M.B. (2020)

New record of Ferrisa dasylirii (Cockerell) (Hemiptera:

Coccomorpha: Pseudococcidae) in Indonesia. Serangga,

(3), 93‒100.

Zarkani A, Sunardi T, Nadrawati, Ercan C, Kaydan MB

(2021a) First record of Rastrococcus tropicasiaticus

Williams (Hemiptera: Coccomorpha: Pseudococcidae)

in Indonesia. Serangga, 26 (3), 29‒36.

Zarkani A, Apriyanto D, Turanli F, Ercan C, Kaydan MB

(2021b) A check list of the scale insect (Hemiptera:

Pseudococcidae) in Indonesia. Zootaxa, 5016 (2), 151‒

Zarkani A, Apriyanto D, Djamilah, Nadrawati, Priyatiningsih,

Sunardi, T., Cansu E., Kaydan M.B (2022) Planococcus

bendovi Williams (Hemiptera: Coccomorpha:

Pseudococcidae), a First Record of Mealybug in

Southeast Asia. Serangga, 27(1): 1-11.

Zarkani A., Ercan C., Apriyanto D., & Kaydan M.B. (2023a).

Studies on mealybugs (Hemiptera: Coccomorpha) in

Indonesia, with description of a new species and three

new country records. Zootaxa 5228(2): 157–172.

https://doi.org/10.11646/zootaxa.5228.2.4

Zarkani A., Fauzi A., Apriyanto D., & Kaydan M.B. (2023b)

Mealybugs (Hemiptera, Coccomorpha, Pseudococcidae)

on parasitic plants (Loranthaceae) in Indonesia with

description of a new species and a new country record.

ZooKeys 1167: 199–210. https://doi.org/10.3897/

zookeys.1167.106012

Zarkani, A., Febrianti, K., Djamilah, D., Nadrawati, Fauzi, A.,

& Apriyanto, D. (2023c) Mealybugs complex of citrus in

Bengkulu Province. E3S Web of Conferences 373

doi.org/10.1051/e3sconf/202337307001

Zarkani A., Fauzi A., Apriyanto D., & Kaydan M.B. (2024a) A

New Planococcus Ferris (Hemiptera: Coccomorpha:

Pseudococcidae) from Indonesia with an Identification

Key of Southern Asia. Journal of Insect Biodiversity and

Systematics 10 (2): 231–242.

Zarkani A., Watson G.W., & Kaydan M.B. (2024b) A new

species in the mealybug genus Pseudococcus Westwood

(Hemiptera: Coccomorpha: Pseudococcidae) from

Indonesia. Zootaxa (in press).

Published

November 22, 2024